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That's a good question. There will be the minimum amount of fluid in the tank when the roof is up and the tonneau and flap are fully extended open. The only difference between minimum and maximum is the displacement of the cylinder rods since there is fluid on both sides of the pistons. This is why it is so difficult to manually actuate the roof since you have to move all that fluid.
Regarding another question, the quarter panel cylinders can be accessed by removing the latch panels on either side of the roof compartment. You can see the hoses go through these panels. If you take them off, you need to make sure they go back into the same location so that the tonneau will latch and fit properly.
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